Main Publications:




1. Chen J, Callac P, Parra LA*, Karunarathna SC, He MQ, Moinard M, Kesel AD, Raspé O, Wisitrassameewong K, Hyde KD, Zhao RL*. 2017. Study in Agaricus subg. Minores and allied clades reveals a new American subgenus and contrasting phylogenetic patterns in Europe and Greater Mekong Subregion. Persoonia 38:170-196.

2. Zhao RL*, Li GJ, Sánchez-Ramírez S, Stata M, Yang ZL, Wu G, Dai YC, He SH, Cui BK, Zhou JL, Wu F, He MQ, Moncalvo JM, Hyde KD. 2017. A six-gene phylogenetic overview of Basidiomycota and allied phyla with estimated divergence times of higher taxa and a phyloproteomics perspective. Fungal Diversity 84(1): 43-74 DOI 10.1007/s13225-017-0381-5

3. Hyde KD, Maharachchikumbura SSN, Hongsanan S, Samarakoon MC, Lücking R, Pem D, Harishchandra D, Jeewon R, Zhao RL, Xu JC, Liu JK, Al-Sadi AM, Bahkali AH, Elgorban AM. 2017. The ranking of fungi: a tribute to David L. Hawksworth on his 70th birthday. Fungal Diversity 84:1–23 DOI 10.1007/s13225-017-0383-3

4. He MQ, Chen J, Zhou JL, Ratchadawan C, Hyde KD, Zhao RL*. 2017. Tropic origins, a dispersal model for saprotrophic mushrooms in Agaricus section Minores with a description of sixteen new species. Scientific Reports DOI: 10.1038/s41598-017-05203-5

5. Chakraborty D, Das K*, BaghelaA A, Mehta N, Singh SK, Mukherjee SK and Zhao RL. 2017. Morphological and phylogenetic studies of Lycoperdon rupicola: first report for the Indian mycobiota. Taiwania 62(1): 93?98. DOI: 10.6165/tai.2017.62.93






1.   Zhao RL*, Zhou JL, Chen J, Margaritescu S, Sánchez-Ramírez S, Hyde KD, Callac P, Parra LA, Li GJ, Moncalvo J-M*, 2016. Towards standardizing taxonomic ranks using divergence times - a case study for reconstruction of the Agaricus taxonomic system. Fungal Diversity 78: 239–292. DOI: 10.1007/s13225-016-0357-x.

2.   Li GJ, Kevin D. Hyde, Zhao RL*, et. al., 2016. Fungal diversity notes 253–366: taxonomic and phylogenetic contributions to fungal species. Fungal Diversity 78: 1–237. DOI: 10.1007/s13225-016-0366-9.

3.   Zhou JL, Su SY, Su HY, Wang B, Callac P, Guinberteau J, Hyde KD, Zhao RL*. 2016. A description of eleven new species of Agaricus sections Xanthodermatei and Hondenses collected from Tibet and the surrounding areas. Phytotaxa 257(2): 99–121. DOI: 10.11646/phytotaxa.257.2.1

4.   Chen J, Parra LA, Kesel AD, Khalid AN, Qasim T, Ashraf A, Bahkali AH, Hyde KD, Zhao RL*, Callac P*. 2016. Inter- and intra-specific variability in Agaricus endoxanthus and allied species reveals a new taxon, A. punjabensis. Phytotaxa 252: 1–16. DOI: 10.11646/phytotaxa.252.1.1

5.   Karunarathna S, Mortimer PE, Chen J, Li GJ, He MQ, Xu JC, Seelan JSSS, Hassan BA, Hyde KD, Zhao RL*. 2016. Correct names of two cultivated mushrooms from the genus Pleurotus in China. Phytotaxa 260: 36–46. DOI: 10.11646/phytotaxa.260.1.4

6.   Chen J, Moinard M, Xu JP, Wang SX, Foulongne-Oriol M, Zhao RL, Hyde KD, Callac P*. 2016 Genetic Analyses of the Internal Transcribed Spacer Sequences Suggest Introgression and Duplication in the Medicinal Mushroom Agaricus subrufescens. Plos ONE 11(5):e0156250. DOI:10.1371/journal.pone.0156250

7.   Karunarathna SC, Chen J, Mortimer PE, Xu JC, Zhao RL, Callac P* and Hyde KD. 2016. Mycosphere Essay 8: A review of genus Agaricus in tropical and humid subtropical regions of Asia. Mycosphere 7 (4): 417–439. DOI: 10.5943/mycosphere/7/4/3

8.   Dai RC, Li GJ, He MQ, Liu RL, Ling ZL, Wu JR & Zhao RL*. 2016. Characterization of four species including one new species of Agaricus subgenus Spissicaules from Eastern China. Mycosphere 7 (4): 405–416. DOI: 10.5943/mycosphere/7/4/2

9.    Xu ML, Li GJ, Zhou JL, Bai XM, Zhao RL*. 2016. New species of Cystolepiota from China. Mycology. DOI:10.1080/21501203.2016.1239231

10.  Parra LA, Tan Y, Xu ML, Zhou JL, Wang B, Zhao RL*. 2016. A reexamination of Allopsalliota indicates synonymy with Micropsalliota (Agariceae, Agaricaceae, Agaricales, Basidiomycota). Mycoscience 57:303–310. DOI:10.1016/j.myc.2016.02.005

11.  Das D, Hembrom M, Parihar A, Zhao RL. 2016. A new species of Cyathus (Agaricaceae) from India. Turkish Journal of Botany 40(1): 97–103. DOI: 10.3906/bot-1311-38

12.  Hongsanan S, Sánchez-Ramírez S, Crous PW, Ariyawansa HA, Zhao RL and Hyde KD. 2016. The evolution of fungal epiphytes. Mycosphere 7 (10): 1538–1561.  Doi 10.5943/mycosphere/7/10/6

13.  Thongbai B, Tulloss RE, Miller SL, Hyde KD, Chen J, Zhao RL, Raspé O*. 2016. A new species and four new records of Amanita (Amanitaceae; Basidiomycota) from Northern Thailand. Phytotaxa 286 (4): 211–231. DOI: 10.11646/phytotaxa.286.4.1

14.   Liu NG, Ariyawansa HA, Hyde KD, Maharachchikumbura SSN, Zhao RL, Phillips AJL, Jayawardena RS, Thambugala KM, Dissanayake AJ, Wijayawardene NN, Liu JK, Liu ZY*, Jeewon R, Jones EBG and Jumpathong J. 2016. Perspectives into the value of genera, families and orders in classification. Mycosphere 7 (10): 1499–1517. Doi 10.5943/mycosphere/7/10/3

15.  He MQ, Chen J and Zhao RL*. 2016. Two new records of Agaricus from Southwest China. Mycotaxon 131:871-880.






1.   Wei L, Li YH, Hyde KD, Zhao RL*, 2015. Micropsalliota pseudoglobocystis, a new species from China. Mycotaxon 130: 555–561. DOI: 10.5248/130.555

Chen J, Hyde DH, Bahkali AH, Zhao RL*, 2015. Micropsalliota brunneosquamuta, a new species from Thailand. Chiang Mai Journal of Science. 2016; 43(X): 1–6 

3.      Chen J, Zhao RL*, Parra LA, Guelly AK, Kesek AD, Rapior S, Hyde KD, Chukeatirote E, Callac P, 2015. Agaricus section Brunneopicti: a phylogenetic reconstruction with descriptions of four new taxa. Phytotaxa 192(3): 145–168. DOI: 10.11646/phytotaxa.192.3.2

4.      Liu JK, Hyde KD, E. B. Gareth Jones, … Zhao RL et. al., 2015. Fungal diversity notes 1–110: taxonomic and phylogenetic contributions to fungal species. Fungal Diversity 72: 1–797. DOI: 10.1007/s13225-015-0324-y.

5.      Ariyawansa HA, Hyde KD, Subashini C. Jayasiri,… Zhao RL et. al., 2015. Fungal diversity notes 111–246: taxonomic and phylogenetic contributions to fungal species. Fungal Diversity 75: 27–274. DOI: 10.1007/s13225-015-0346-5 (SCI Journal Position 6.9%)

6.      Jayasiri SC, Hyde KD, Ariyawansa HA, .., Zhao RL, al. 2015. The Faces of Fungi database: Fungal names linked with morphology, molecular and human attributes. Fungal Diversity 74: 3–18. DOI: 10.1007/s13225-015-0351-8 (SCI Journal Position 6.9%)

7.      He MQ, Zhao RL*. 2015. A new species of Agaricus section Minores from China. Mycology 6: 182–186.  DOI: 10.1080/21501203.2015.1121931

8.      Wang ZR, Parra LA, Calla P, Zhou JL, Fu WJ, Dui SH, Hyde KD, Zhao RL*. 2015. Edible species of Agaricus (Agaricaceae) from Xinjiang Province (Western China). Phytotaxa 202 (3): 185–197. DOI: 10.11646/phytotaxa.202.3.2


Before 2014


1.      Zhao RL* , Desjardin ED, Callac P, Parra LA, Guinbereau J, Soytong K, Karunarathna SC, Zhang Y, Hyde KD. 2013. Two species of Agaricus sect. Xanthodermatei from Thailand. Mycotaxon 122: 187–195

2.      Das K, Zhao RL* . 2013. Nidula shingbaensis sp. nov., a new bird’s nest fungus from India. Mycotaxon 125: 53–58

3.      Chen J, Zhao RL*, Karunarathna SC, Callac P, Raspé O, Bahkali AH, Hyde KD2012.  Agaricus megalosporus: a new species in section Minores. Cryptogamie, Mycologie, 33 (2): 145–155

4.      Zhao RL, Hyde KD, Desjardin DE, Raspe′ O, Soytong K, Guinberteau J, Karunarathna SC, Callac P. 2012. Agaricus flocculosipes sp. nov., a new potentially cultivatable species from the palaeotropics. Mycoscience 53 (4): 300–311

5.      Karunarathna SC, Yang ZL, Zhao RL, Vellinga EC, Bahkali AH, Chukeatirote E, Hyde KD. 2011. Three new species of Lentinus from northern Thailand. Mycological Progress 10 (3): 389–398

6.      Zhao RL, Karunarathna S, Raspé O, Parra LA, Guinberteau J, Moinard M, Kesel AD, Barroso G, Courtecuisse R, Hyde KD, Guelly AK, Desjardin ED, Callac P. 2011. Major clades in tropical Agaricus. Fungal Diversity 51:279–296

7.      Karunarathna SC, Yang ZL, Raspe O Ko Ko TW, Vellinga EC, Zhao RL, Bahkali AH, Chukeatirote E, Degreef J, Callac P, Hyde KD. 2011. Lentinus giganteus revisited: new collections from Sri Lanka and Thailand. Mycotaxon 118:57–71

8.      Thongklang N, Nawaz R, Khalid AN, Chen J, Hyde KD, Zhao RL, Parra LA, Hanif M, Moinard M, Callac P. 2014. Morphological and molecular characterization of three Agaricus species from tropical Asia (Pakistan, Thailand) reveals a new group in section Xanthodermatei. Mycologia 106(6): 1220–1232.

9.      Sysouphanthong P, Thongkantha S, Zhao RL, Soytong K, Hyde KD. 2010. Mushroom diversity in sustainable shade tea forest and the effect of ?re damage, Biodiversity and Conservation 19 (5): 1401–1415

10.  Zhao RL, Desjardin DE, Soytong K, Perry BA, Hyde KD. 2010. A monograph of Micropsalliota in Northern Thailand based on morphological and molecular data. Fungal Diversity vol.45: 33–79

11.  Zhao RL, Desjardin DE, Soytong K, Hyde KD. 2008. A new species of bird's nest fungi: Characterization of Cyathus subglobisporus sp. nov. based on morphological and molecular data. Persoonia 21: 71–76.

12.  Zhao RL, Jeewon R, Desjardin DE, Soytong K, Hyde KD. 2007. Ribosomal DNA phylogenies of Cyathus: Is the current infrageneric classification appropriate? Mycologia 99(3):385–395.

13.  Zhou TX, Zhao LZ, Zhao RL* and Chen YH. 2004. The Notes on the Bird’s Nest Fungi from China.  Fungal Diversity. 17: 243–251


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